Celebration Potato Pancakes Recipe, Spotlight on Smoked Salmon, Benefits of a Hot Pan, and Flavonoids’ Mood-Boosting Benefits
A wonderful late autumn dish, potato pancakes with smoked salmon are tasty and satisfying. And you can elevate them from everyday to festive with flavorful garnishes perfect for the most elegant celebrations. I’m also sharing welcome news about the mental health benefits of foods rich in flavonoids and an invitation to sign up for a bone health masterclass from Kevin Ellis, the Bone Coach—the time is now to protect your bones.
This twist on traditional potato pancakes gets sweetness from parsnips and a hint of tartness from the apple, plus garnishes that add more levels of flavor. Eggs replace flour as the binder, making this version gluten free as well.
Ingredients
12 ounces red potatoes, scrubbed but not peeled
1 medium yellow onion, about 8 ounces
1 large Granny Smith apple, washed
6 ounces parsnips, peeled
3 large eggs
4 tablespoons extra virgin olive oil, divided use, plus more as needed
2 teaspoons coarse sea salt
1 teaspoon freshly ground black pepper
1 cup skyr, sour cream, or mascarpone
4 ounces thinly sliced smoked salmon, cut into 32 pieces
A few springs of fresh dill
Additional garnishes: capers, salmon roe or another caviar, diced red onion
Directions
Step 1
Using the grating blade of a food processor or a large box grater, finely grate the potatoes, onion, apple, and parsnips. Transfer to a very large bowl and mix well. Add the eggs, the 2 tablespoons olive oil, salt, and pepper, and toss thoroughly. Preheat your oven to 250°F and line a rimmed sheet pan with paper towels.
Step 2
Heat a large frying pan over medium heat. When hot, add 2 tablespoons of olive oil. Using a 1/4 cup measure as a scoop, make a layer of pancakes, flattening them slightly with the back of the measure. Cook over medium heat until they brown on the bottom, flip, and continue cooking until the underside browns and crisps. Transfer to the sheet pan and place in the oven to keep warm. Repeat with the rest of the mixture, adding more oil to the pan as needed.
Step 3
To serve, top each pancake with a dollop of skyr, sour cream, or mascarpone; a piece of smoked salmon; a snippet of dill; and a few capers, roe, or another caviar, if desired.
Yields 8 servings
Healthy Ingredient Spotlight
Smoked Salmon
Whether you prefer your smoked salmon Scandinavian style with butter on dark bread or New York style with cream cheese on a bagel, this specialty fish has the same health benefits as grilled or poached salmon plus a salty, smoky flavor. It retains salmon’s natural omega-3 fatty acids, which help boost the health of your heart, brain, and eyes.
Smoked salmon also delivers on protein, plus vitamins A, B12, and E and the mineral selenium, all for about 30 calories per ounce. Do read labels to make sure that the sodium content from the curing process won’t put you over your limit—its salty taste means you can skip the salt shaker when eating it as a topper on your potato pancakes or avocado toast, and when adding it to scrambled eggs.
Quick Kitchen Nugget
Start with a Hot Pan
I’m not a fan of adding olive oil to a cold pan—that has the effect of cooking the oil while the pan itself gets to the right temperature. Instead, heat your pan or pot over medium-high heat for a couple of minutes; when a few drops of water sprinkled on the surface sizzle, it’s time to add your EVOO and then the food to be cooked.
For Your Best Health
Flavonoids’ Mood-Boosting Benefits
According to research presented at the annual meeting of the American Society of Nutrition by Department of Veterans Affairs researcher Galya Bigman, PhD, having a lot of flavonoids in your daily diet is linked to a reduced risk of depression. Flavonoids are a key group of natural compounds found in plant foods and offer a variety of health benefits. For this work, Dr. Bigman used data from the National Health and Nutrition Examination Survey (NHANES: 2007–2010, 2017–2018) including self-reported depressive symptoms and dietary intake of flavonoids from more than 12,000 participants. Total flavonoid intake (mg/day) was calculated and divided into quartiles, with participants’ flavonoid intake was ranked.
Participants in the highest vs. lowest quartiles of flavonoid intake had a 10 to 13 percent lower incident rate of depressive symptoms, leading to the conclusion that “flavonoids may have a central role in alleviating or preventing depressive symptoms.” While further research is needed to clarify the mechanisms responsible for this positive link, the research notes the top food sources of flavonoids that the participants enjoyed: cocoa/chocolate, tea, soy-based products, onions, chili/sweet peppers, berries, dark-green leafy vegetables, celery, and lemon.
Roasted Potatoes and Foolproof Scrambled Eggs Recipes, Spotlight on Urfa Chili and Oregano Pearls, Measuring Spices, Plus Spices and Your Health
The saying goes that, for better health, eat dinner like a pauper and breakfast like a king or queen. There are even research studies to support the idea: because breakfast prompts increased calorie burn, it can be helpful for staying at or losing weight. Of course, we don’t need science to tell us that breakfast can be delicious, too, and I’m sharing two of my favorite recipes to prove it. They feature spices that I like to source from the magical land of Turkey—oregano and black Urfa chili—and they truly elevate this classic eggs-and-potatoes meal.
Fragrant and richly hued, yellow rice takes the standard side dish and elevates it. This recipe gets another taste boost from the green peas.
Ingredients
2 pounds red or Yukon gold potatoes, scrubbed and cut into roughly 2-inch cubes
2 tablespoons extra virgin olive oil
1 teaspoon dried oregano
1/2 teaspoon Black Urfa Chili
1 teaspoon coarse sea salt
Directions
Place a rack in the middle of your oven and preheat to 450°F. In a large bowl, toss the potatoes with the oil, spices, and salt. Roast for 50 minutes, flipping the potatoes and rotating the pan after 25 minutes. Taste and season with more spices as desired.
A properly preheated frying pan makes fast work of scrambled eggs that stay tender. Black Urfa chili adds just the right amount of zest to the finished dish while the oregano adds brightness.
Ingredients
8 large or extra-large eggs
2 tablespoons extra virgin olive oil, plus more for drizzling
Black Urfa Chili and dried oregano
Directions
Whisk the eggs in a bowl. Heat a frying pan over medium-high heat. When the pan is hot, add the olive oil and then the beaten eggs. Let the edges firm for a few seconds, then lower the heat and use a spatula to scramble the eggs, drawing the outer edges in and keeping them moving to prevent browning. Take off the heat just before fully set and let sit for two minutes to finish cooking. Season to taste with the black Urfa chili and oregano, then drizzle with olive oil.
Yields 4 servings
Healthy Ingredient Spotlight
Urfa Chili and Oregano
Black Urfa Chili
Black Urfa Chili (Capsicum annuum) takes its name from the ancient southeast town of Urfa, Turkey, where local farmers grow it in the surrounding hills. In many areas of that country, Urfa chili, also known as Urfa biber, is as common on the dinner table as cracked black peppercorns are here, and it has been for centuries. These peppers are red and shaped like bell peppers, but they pack a taste that a bell pepper can only dream of!
Urfa peppers magically turn maroon as they ripen. Then, unlike most peppers that are simply dried after harvest, they go through a double process—sun dried by day and tightly wrapped at night, a technique that preserves their oil content, enriches their flavor, and deepens their color to a purple-black. They’re then stone ground into small flakes with some sea salt and sunflower seed oil to further enhance their texture and rich, complex flavor profile—the deep heat of cayenne without the sharpness.
Why you should have this spice in your kitchen: Black Urfa chili adds sweet smokiness to all Middle Eastern dishes, and it’s especially delicious on boreks,koftis, kebabs, and other grilled meats, and on veggies. It’s equally good in many other cuisines—think French stew or an all-American chili. Include it in wet or dry rubs for proteins, especially pork. It will enliven your favorite salad dressings, jazz up scrambled or fried eggs, and add extra depth and a hint of heat to brownies, blondies, hot chocolate, and even a scoop of chocolate ice cream.
Oregano Pearls
Oregano’s popularity among American cooks is relatively recent. Legend has it that GIs returning from World War II brought back their new love of pizza, liberally sprinkled with the herb, and its use here exploded. Though closely associated with classic dishes of southern Italy, oregano (Origanum vulgare) is an essential in cuisines throughout the Mediterranean region. It’s also popular in Latin America as well as Mexico, and it is the perfect herb for chile-based dishes. But know that what’s labeled “Mexican oregano” is actually a member of the Verbenaceae family and not a true oregano (it’s also much sharper).
Oregano was first cultivated in Greece. Mine comes from Denizli, Turkey, located in the heart of the Aegean region, just a few hours’ drive from the sea. The country is known for growing prized oregano—indeed, Turkey satisfies more than half of global demand. Dried oregano is typically made from the leaves, but my Curated Culinary Selections’ oregano “pearls” (complete with a grinder) are the tiny buds of the oregano flowers. And timing is everything: They’re handpicked before they have a chance to bloom and then are air dried. This special process gives them a bright flavor and makes them perfect for grinding as needed.
Why you should have this spice in your kitchen: Perfect in tomato- and meat-based sauces and bold pasta dishes, oregano is also wonderful when paired with garlic—for the best garlic bread, brush slices with fresh-pressed extra virgin olive oil and top with a few grinds of oregano pearls before they go into the oven. Flat breads and focaccia will benefit, too. Oregano adds great flavor to grilled fish, roasted chicken, and all kinds of meat, especially lamb (kebabs and patties in particular), and to your favorite chili recipe. Beyond pasta and pizza, sprinkle it on Greek salad,omelets, frittatas, roasted potatoes, and even fries.
For Your Best Health
Spices and Your Health
As with all chile peppers, at the heart of black Urfa chili’s heat is capsaicin, the naturally occurring compound that determines how sweet or hot a pepper is. Capsaicin has antioxidant and antibacterial properties, so it may help protect against infection, improve digestion, and possibly one day have a role in anti-cancer therapies. According to a report published in the journal Molecules, there’s some evidence that eating hot pepper, like Urfa chili, every day helps curb hunger and may even lead to a higher calorie burn rate. Research is underway to figure out the best daily amount, but in the meantime you can test out the theory yourself.
Oregano is one of the most widely used botanicals in herbal medicine, thanks to its high concentration of plant nutrients. Phytochemical compounds such as flavonoids and phenolic acids give it the triumvirate of health benefits: antibacterial, antioxidant, and anti-inflammatory properties. Sipping a tea made from dried oregano, for instance, can calm an upset stomach or a cough. It’s also being looked at to help stave off diabetes—researchers at the University of Illinois found that it disrupts a diabetes-related enzyme and merits serious investigation. It may also boost liver health.
Quick Kitchen Nugget
Measuring Out Spices
While measuring spoons are the tried-and-true way most people measure quantities, you might find more and more references to gram weights for ingredients in recipes. That’s because weight can be more precise than volume, especially when measuring whole spices that don’t fit perfectly in a spoon. An electronic scale with a bright digital readout will be a great addition to your kitchen gadgets.
Yellow Rice and Green Beans with Pimentón Recipes, Spotlight on Pimentón and Turmeric, Storing Spices and Your Health, and How Restricting Calories Can Boost Longevity
Want to elevate your cooking this holiday season? Grab your spices and turn dull into delicious! In this installment of my single-origin spice series, I’m focusing on two high-flavor choices that may seem exotic, yet can be used often and with tempting results.
Remember too that spices aren’t just for cooking—many are great for boosting the flavor of finished dishes. So, bring some out when you set your table. You and your guests won’t reach for the salt shaker as often, and that’s good for your health.
Fragrant and richly hued, yellow rice takes the standard side dish and elevates it. This recipe gets another taste boost from the green peas.
Ingredients
1 tablespoon extra virgin olive oil
1 tablespoon unsalted butter
1/4 cup minced onion
1 cup basmati or jasmine rice
1/2 teaspoon turmeric
1-1/2 cups water or chicken broth, homemade or low-sodium store bought
4 strands saffron, optional
1 cup frozen green peas, thawed
Directions
Heat a 2-quart saucepan over medium heat. When hot, add the olive oil and butter, then the onions. Cook, stirring frequently, until softened but not browned, about 3 minutes. Add the rice and turmeric, stirring to coat the grains. Add the water or broth and saffron, if using, and bring to a low boil. Reduce heat to a simmer, cover, and cook for 20 minutes, checking after 15 to see if all the liquid has been absorbed. When done, turn off the heat, fluff with a fork, fold in the peas, and cover the saucepan for 2 minutes before serving.
Wonderful when made with thin French green beans called haricots verts, often available frozen year-round.
Ingredients
1-1/2 pounds fresh or frozen green beans
4 to 6 tablespoons extra virgin olive oil
2 red bell peppers, stemmed, seeded, and cut into strips
3 cloves garlic, coarsely chopped
1 teaspoon Pimentón or more to taste
3 ounces cured ham, preferably Serrano, cut into 1/4-inch cubes
Freshly ground black pepper, to taste
Coarse sea salt, if needed
Directions
If using fresh beans, trim the ends and steam them until tender-crisp, about 5 minutes. Immediately plunge them into a large bowl of ice water to stop the cooking and set the color. Drain and set aside. Heat a large frying pan over medium heat. When hot, add 2 tablespoons of olive oil and the bell peppers, sautéing the peppers until softened, 4 to 6 minutes. Add the garlic and sauté for 1 minute. Add 2 more tablespoons of olive oil and the pimentón, stirring well to distribute the spice. Add the ham and the steamed beans. Season with black pepper and, if needed, salt (the ham itself may add enough salt), and stir the beans with tongs to heat through. If they seem dry, add the rest of the oil. When the beans are ready, taste and add more pimentón, black pepper, and/or salt as desired.
Yields 4 servings
Healthy Ingredient Spotlight
Pimentón and Turmeric
Pimentón
All peppers were brought to Spain from the Americas, but the peppers used to make pimentón (Capsicum annuum) have a history all their own. They were given to monks at the Yuste monastery in La Vera, an area within the region of Extremadura, by King Ferdinand and Queen Isabella. The monks dried them and turned them into a powder. Peppers later made their way to the rest of Europe, notably Hungary, where the unsmoked version of the powder became that country’s classic spice, paprika.
The distinct sweet smoky aroma and flavor of pimentón come from thelocalclimate and the unique, centuries-old and labor-intensive smoking process. After being handpicked, the peppers go to a two-story smokehouse. On the first story is an oak fire that burns around the clock. The peppers are smoked on the second story—for two weeks, they’re continually bathed in the aromatic heat and regularly rotated for even crispness. The spice’s full name is Pimentón de La Vera DOP. DOP means “Designation of Protected Origin,” a label bestowed when a specific region is recognized as producing certain foods with special local characteristics. It’s pimentón that, in turn, gives the region’s famed chorizo and other pork products their distinctive flavor and color.
Why you should have this spice in your kitchen: Pimentón defines many Spanish dishes, such as shrimp with pimentón and garlic,patatas bravas, and paella. It’s also wonderful in deviled eggs, dry rubs, and marinades for barbecues, and, for the adventurous, homemade sausages. It marries perfectly with fresh-pressed extra virgin olive oil to add heat to any sauté or to popcorn made on the stovetop. I love it on braised greens and cooked legumes.
Golden Turmeric
India is the land of intriguing spices, rich and deep, and none more colorful than turmeric (Curcuma longa). It belongs to the Zingiberaceae family, which also includes ginger, and you can see the similarity between them when you hold up the edible parts, the rhizomes. Also, like ginger, turmeric has been used for thousands of years, and not only in India but throughout South Asia and into the Middle East, Africa, and beyond, in both sweet and savory dishes. Besides its culinary tradition, turmeric is used in Eastern medicine, in religious ceremonies, and even as a natural coloring.
Why you should have this spice in your kitchen: Besides using turmeric in your favorite Indian dishes from kormas to dals, add it to sauces, stews, broths, salad dressings, juices, and baked goods for a depth of flavor and a jolt of color. Toss cut-up root vegetables with olive oil, sprinkle with turmeric, and roast at 400°F until soft. For a soothing brew, createa paste with turmeric, honey, cinnamon,and a healthy pinch of freshly ground black peppercorns, and whisk it into a cup of warmed milk of your choice.
Quick Kitchen Nugget
Storing Spices
The best way to store spices isn’t the handiest—they should be in tightly sealed glass containers away from light, humidity, and heat, so not above the cooktop. Keep them on a dry pantry shelf or in a drawer away from the oven, and soon it will be second nature to take them out of their new “home” as you prep ingredients and set the table with these great alternatives to salt.
For Your Best Health
Spices and Your Health
Peppers have among the highest levels of antioxidants of any vegetable, including bioactive compounds like vitamin C, phenolics, and carotenoids (the source of their color), all strong disease fighters. The carotenoids include zeaxanthin and lutein, good for eye health, according to International Journal of Molecular Sciences. Pimentón’s spiciness comes from capsaicin, a compound with anti-inflammatory properties. While the amount of all this goodness in a little pinch of the spice is small, all those little pinches can add up to help meet micronutrient needs.
Most of the interest in turmeric comes from its powerful polyphenol curcumin, thought to have anti-inflammatory and antioxidant properties, though research done in Western countries hasn’t quite been able to explain how it works. A newer concept, published in Advances in Nutrition, is that curcumin could have a direct positive effect on the GI tract and that its health benefits could emanate from there. Interestingly, we know that, on its own, curcumin has a low bioavailability, meaning the body isn’t able to draw out and make the most of its nutrients. According to a report in the journal Foods, you can increase that bioavailability by having it along with black pepper, rich in the active ingredient piperine. So, every time you reach for turmeric, grab your pepper corn grinder, too.
Guacamole and Pico de Gallo Recipes, Spotlight on Cumin and Black Lime, Toasting Spices and Your Health, and a 6th Sense Discovered!
Let’s get the party started! Who doesn’t love guac and salsa? (My pico de gallo recipe, below, is one of the simplest salsas you can make.) Just add your favorite dippers—I love bell pepper spears for the extra nutrients—and you’ll have a happy crowd.
Both of this issue’s recipes feature two of the most flavorful spices on the planet: wild cumin and black lime (read on to discover more about them). If these are new to you, you’ll be pleasantly surprised by their depth of flavor. I love them both but was frustrated by not being able to find them easily in stores. So, I became determined to locate harvest-fresh, single-origin spices. The latest spice collection in my Curated Culinary Selections includes them both along with six other spice essentials.
Speaking of taste sensations, I’m also sharing the first taste bud discovery in over a century—turns out we have a sixth taste. I’m sure you’ll find the news as interesting as I did!
This guacamole gets a jolt of flavor from cumin. You can intensify the heat by leaving in some of the jalapeño seeds.
Ingredients
2 garlic cloves, peeled
1 small jalapeño
1 small onion
1 cup of cilantro, leaves and some stems
3 large ripe avocados
1 tablespoon extra virgin olive oil, or more to taste
1 lime, halved
1 teaspoon cumin (wild if possible), toasted and ground
Freshly ground black peppercorns
Coarse sea salt
Directions
Mince the garlic and set it aside for 10 minutes to allow its healthful compound allicin to develop. Wearing gloves to keep the jalapeño’s powerful oils off your skin, slice the pepper in half lengthwise and use a small spoon to scrape out the ribs and seeds, and then cut it into a small dice. On a large cutting board, chop the onion into a large dice and keep chopping as you add in the garlic, jalapeños, and cilantro to meld them together. Halve the avocados and use a spoon to scoop all the flesh into a large bowl. Add the olive oil, coarsely mash the avocado with a fork, and then fold in the onion mixture. Squeeze in the juice of a lime half and fold in the cumin, a few twists of your peppercorn grinder, and a sprinkle of salt. Taste and adjust the seasonings, adding more olive oil, lime juice, black pepper, and/or salt as desired.
I love this Mexican condiment for its bold flavors and easy prep—no machines needed, just a good sharp knife to dice the main ingredients. Besides being a delicious dip for chips and veggies, it’s a great condiment for grilled chicken and fish for your next Taco Tuesday.
Ingredients
2 garlic cloves
2 large fresh tomatoes or 6 canned Roma tomatoes
1 medium onion
1 jalapeño or other chile pepper
1/2 cup chopped cilantro (leaves and some stems)
1 teaspoon cumin seeds, toasted and ground
Juice of 1/2 lime
Freshly ground black pepper
Coarse sea salt
Black Lime
Directions
Mince the garlic and set aside. Dice the tomatoes and onion. Wearing gloves to keep the powerful oils off your skin, slice the jalapeño in half lengthwise and use a small spoon to scrape out the ribs and seeds, and then cut it into a small dice. Place the vegetables in a large bowl along with the garlic and cilantro, and toss well. Fold in the cumin and lime juice. Taste, add a few twists of the black peppercorn grinder and a sprinkle of the salt, and taste again, adding more salt and pepper as desired. Let the flavors meld at room temperature for 30 minutes and then refrigerate until serving. Just before serving, top with a sprinkle of black lime.
Yields about 2 cups
Healthy Ingredient Spotlight
Cumin and Black Lime
Foraged Wild Cumin
Cumin is used in so many cuisines, not only those of countries where it was first grown but also in others a world away. The common variety (Cuminum cyminum) is indigenous to the Middle East and central regions of India. My preference, wild cumin (Bunium persicum), used in cooking for centuries, has a somewhat mysterious history because it’s native to very remote areas of the world. These include regions of central Asia such as Iran, Pakistan, and the Himalayas in northern India, where it is known as kala jeera—you may see it listed that way in recipes, including Mughlai cuisine. The small, dark-brown seeds of wild cumin are extremely rare. They’re not farmed but rather handpicked from wild plants and then sun dried.
Why you should have this spice in your kitchen: Wild cumin, used whole or ground, shines in classic Indian dishes such as biryanis and curries, Middle Eastern falafel and hummus, and North African tagines and harissa, a heavily spiced chile paste. Cumin also is key to many exotic spice blends, such asbaharat and dukkah. But don’t be shy about using it in Mexican and Tex-Mex dishes that typically call for plain cumin, in beans and chili, and in guacamole and pico de gallo.It also adds wonderful zest when sprinkled whole on naan and other flatbreads and when ground for dry rubs and marinades.
Black Lime
Black lime has a rich and storied legacy in Persian cuisine, dating back centuries, and has been used for everything from making a delicious tea to dyeing yarn. It’s popular in northern India, in Iraq, and in the Arab states of the Persian Gulf, including Saudi Arabia, Kuwait, and Oman, where it’s known as loomi or limu Omani. Still, many American cooks have never used black lime before, and it’s hard to find outside of some ethnic food stores. To create black lime,ripe limes are dried in the sun to naturally turn black. While black limes are added whole in many Persian recipes, the ground version is very handy to use, plus the grinding process releases their fragrant oils to the max.
Why you should have this spice in your kitchen: Black lime is the brightest and most intense lime zest you’ve ever tasted. It’s especially delicious on seafood—grilled shrimp, salmon, and tuna—and on meat, rice and other grains, and vegetables. Try it on ceviche, avocado toast, and soft cheeses; as an ingredient in vinaigrettes; in marinades, dry rubs, and za’atar, a spice blend especially delicious with for lamb and chicken; and on any food you like with a squeeze of lime. A fun way to enjoy its flavor jolt is to use it to rim the glasses the next time you mix up a batch of margaritas or bloody Mary’s.
Quick Kitchen Nugget
Toasting Whole Spices
Lightly toasting whole spices like cumin seeds intensifies their flavor. The easiest way is in a small frying pan on the stovetop over medium heat. It’s fast—about three minutes—but requires your attention for the entire time because they can burn in a snap. Shake the pan every few seconds to rotate the spices. As soon as you smell the aroma rising from the pan, they’re done. Let cool briefly before grinding or proceeding with a recipe.
For Your Best Health
Spices and Your Health
Wild cumin has a bounty of antimicrobial, anti-inflammatory, and antioxidant properties. Many cultures have used the essential oils pressed from the seeds as a botanical remedy for digestive and other problems for centuries. Because different strains grow wild in different parts of the world, it’s hard to study its health benefits in a formal setting to truly unlock its potential, according to research in the journal Physiology and Molecular Biology of Plants.
It’s hard to say how many of the lime’s natural plant compounds, such as vitamin C and other antioxidants, remain intact or, conversely, become intensified by the drying process of making black lime. One thing is certain: It’s a great way to season food without reaching for the salt shaker, and limiting salt has great health benefits, including keeping blood pressure in a safe zone, according to the American Heart Association.
Food Flash
Discovered: A Sixth Taste!
Back in the early 1900s, Japanese scientist Kikunae Ikeda first proposed umami as a basic taste in addition to sweet, sour, salty, and bitter. About eight decades later, the scientific community officially agreed with him. Now scientists led by researchers at the USC Dornsife College of Letters, Arts and Sciences have unraveled the mystery surrounding a sixth basic taste.
It’s been known for some time that the tongue responds strongly to ammonium chloride. However, the specific tongue receptors responsible for it remained elusive until now. In research published in Nature Communications, USC neuroscientist Emily Liman, PhD, and her team found that the response is triggered through the same protein receptors that signal sour taste (the USC team discovered the protein responsible for detecting sour, too).
Called OTOP1, it sits within cell membranes and forms a channel for hydrogen ions moving into the cell. Hydrogen ions are the key component of acids, and as foodies everywhere know, the tongue senses acid as sour. That’s why lemonade (rich in citric and ascorbic acids), vinegar (acetic acid), and other acidic foods impart a zing of tartness when they hit the tongue. Hydrogen ions from these acidic substances move into taste receptor cells through the OTOP1 channel.
Because ammonium chloride can affect the concentration of acid—that is, hydrogen ions—within a cell, the team wondered if it could somehow also trigger OTOP1. To answer this question, they introduced the OTOP1 gene into lab-grown human cells so the cells would produce the OTOP1 receptor protein. They then exposed the cells to acid or to ammonium chloride and measured the responses. “We saw that ammonium chloride is a really strong activator of the OTOP1 channel,” Dr. Liman says. “It activates it as well as or better than acids.
“If you live in a Scandinavian country, you will be familiar with and may like this taste,” says Dr. Liman, who is also a professor of biological sciences. In some northern European countries, salt licorice has been a popular candy for over a century. The treat counts among its ingredients salmiak salt, or ammonium chloride.”